Major Owen Hunt returns to be the head of ER trauma; Meredith finds an old doll her mother gave her when she was 5.
A woman, using her professional skills as a call girl, becomes the team's target as she goes after high-powered executives.
Susan receives some surprising news while she's away on a romantic weekend with Ian, Lynette continues to try to support the family while Tom chases his dreams, and Bree finds her family is anything but perfect.
Jane and the team narrowly escape a fiery end when a building explodes while they are investigating a death threat. Meanwhile, sparks fly between Lisbon and billionaire Walter Mashburn when they meet again during a case.
When an Arctic explorer is killed during a polar expedition, the mysterious circumstances of his death take Castle and Beckett from the penthouses of Park Avenue to the halls of the CIA.