Ed Mercer's enthusiasm about his promotion to captain dampens when his ex-wife, Kelly, gets assigned to be his first officer; an alien race ambushes the crew in an attempt to steal a time-accelerating device.
Ed and Kelly find themselves prisoners in a replica of their former home; Alara must take first command of the Orville to attempt a heroic rescue; Bortus and Klyden receive unexpected news.
The Orville travels to Bortus' planet; the crew debates controversial surgeries for newborns.
Ed and the Orville crew encounter a massive vessel adrift in space, which is in danger of colliding with a star. While some of the crew set out to help the ship’s inhabitants, Bortus, Gordon, and John stay behind and fend off a Krill attack on another Union ship.
After the crew saves a stranded ship and its attractive captain (guest star Charlize Theron), Ed becomes smitten with the Orville’s new guest, while Kelly suspects all is not what it seems. Meanwhile, Gordon attempts to teach Isaac the art of pulling off a practical joke.
After the Orville crew defeats a Krill attack on a brand-new colony, the Union sends Ed and Gordon on a dangerous undercover mission to infiltrate a Krill ship and obtain a copy of the Krill bible, the “Anhkana,” to better understand the enemy.
Seth MacFarlane
Captain Ed Mercer
Adrianne Palicki
Commander Kelly Grayson
Penny Johnson
Dr. Claire Finn
Scott Grimes
Lieutenant Gordon Malloy
Peter Macon
Lieutenant Commander Bortus