Four years passed since Sagan's called a truce with the D2s; music was still taboo throughout the world; somewhere in America, a little boy who grew up not knowing what music is hears the melody from a piano.

D2s attack Boston, a blood-soaked concert hall, the incident known as the Boston Tragedy; the conductor who lost his life in that tragedy was none other than Kenji Asahina, Takt's father; he decides to devote himself to playing the piano.

Cosette transforms into the Musicart named Destiny; Destiny proclaims that her mission is to eliminate the D2s; Destiny bestows a conductor's baton upon Takt, and he uses his Conductor powers for the first time.

Takt and Destiny learn how to fight properly from Lenny and Titan as they continue to make their way to New York.

Takt and crew continue their journey under the blazing sun; during their trip, they end up fighting some D2s and end up bringing a train on the transcontinental railroad to a sudden stop.

Takt and crew stop by a certain town; the residents ask Anna and Destiny for help; Takt wandered around town on his own and happens to hear a melody quietly resonating from an old bar.