In the distant future, the planets are now connected with a massive galactic railway system. The Galaxy Express 999 has crash landed on a mysterious planet. On board are The Conductor, Killian, Maetel, and Tetsuro, a ten year-old street urchin with a heart of gold. Now it's up to the SDF Sirius Platoon to rescue them before a rift in the deep axis of time and space repeats.