Meredith's dishonesty is revealed, leading to unexpected consequences for her and others; Cristina is forced to choose between her relationship and her career; the new chief resident is chosen.
Blake is forced to work with her father, a retired police captain, and brother, a detective, to solve a number of murders.
After the mysterious death of an astronaut in a highly sensitive Mars simulation, the team must send Ernie into the habitat to investigate alone.
Randall and Beth struggle to adjust to the family changes, Kate gets her first big gig, and Kevin bumps into a problem on his movie.
Kate and Toby experience a sudden change in their lives' trajectories, and Rebecca is supportive of Kate's aspirations to become a singer.