Jian Leonardo

Betty, Marc and Matt team up to work on their final YETI project; Daniel proposes to Molly; Matt's mother (Christine Baranski) asks Betty for a favor.

A Mode editor's sudden death leaves Betty and Marc competing for the same job; Wilhelmina demands that Claire resign in exchange for her silence about Claire's dark secret.

Matt asks someone to be his "plus one" at Wilhelmina's fundraiser; Daniel hires Natalie to be his assistant, making Betty feel obsolete; Amanda keeps a secret from Betty; Daniel discovers that his mother has a history with Cal Hartley.

Betty strives for a scoop with a new designer; Marc enlists clueless Amanda and Matt in his effort to sabotage Betty; Daniel attends bereavement sessions, where he meets Natalie (Jamie-Lynn Sigler), who takes him out for the night.

Hilda and Betty have pregnancy scares; Marc has a plan to get Wilhelmina back; Claire finds the son she gave up for adoption.

Daniel and Wilhelmina make Mode's issue based on sex to spice things up. Matt is hesitant to take the next step with Betty. Marc and Amanda provide Betty with a makeover.