Hyuga Saki and Misho Mai meet beneath the Sky Tree, a place that holds special significance to both of them; before they get a chance to talk, they're interrupted by two cute little spirits, who tell them to become Pretty Cure and save the world.

Saki and Mai get to know each other better and Saki decides to set up a welcome party for Mai; but the ultimate party crasher shows up in the form of Karehan.

The Nagi Middle softball team's first match of the season is against the previous year's tournament winners; Saki doesn't think she's intimidated, but as the match starts to go south, Mai notices something crucial.

Mai needs to find a spring subject for an upcoming art contest; she heads out with Saki and they find a field of cabbages; but they hear a sound from the forest.

Saki develops a crush on Mai's brother; she even borrows a complicated book on astronomy from him; when Kazuya spills tea on the pages, Saki's temper hits the roof.

Minori is unhappy that her parents have to run the bakery, so they can't go out on Sundays; Mai's parents offer the girls a trip to a nearby lake.