The alien glyphs Coulson has been etching lead the team to several murder victims; lies and secrets come to a head.
A colleague's death leads Will and Mac to think about the milestones that transformed the newsroom; Maggie lands an interview in Washington; Neal's digital site is shut down for repairs; Leona offers advice to Pruit.
The family is dismayed when John tries to make himself look younger; an over-the-top birthday party and overachieving parents make Jen and Greg worry about getting Lark into a good preschool; Matt meets Colleen's mother.
Tyler's great-grandmother wants to give him her car, since her driver's licence was taken away, but the offer comes with a catch. Jen looks for a creative way to break Greg's habit of falling asleep on the couch instead of coming to bed.