Patrick Aryee follows the grizzly bears, who are taking a risk with the weather by leaving their winter dens early. Hungry wolves are struggling to bring down their elk prey in the unusually shallow snow. And for great grey owls it's the iciness of the snow that is hampering their hunts. Yellowstone’s winter is always one of the most brutal on the planet. But 2016 saw weather records broken, and the wildlife was forced to adapt to survive. Kate Humble gets to grips with the science behind this remarkable season.

As winter turns to spring, temperatures rise and Yellowstone bursts into life.

It’s summer - and the Yellowstone beavers have a new challenge. Will the young survive as the river dries up and the colony is forced to move home? As food becomes scarce wolves have a surprising strategy to keep their pups fed. By midsummer the hot dry conditions create a new danger - deadly wildfires burn out of control and threaten to engulf a family of great grey owls. 2016 was the hottest year on earth since records began, and across Yellowstone scientists reveal the effects of rising temperatures on the animals that live here.