Arriane Alexander

Following a plane crash, the doctors of Seattle Grace try to cope with loss, adapt to changes and move forward with their lives.

Sheldon has a scientific breakthrough at the dentist’s office when he has to get a tooth pulled. Meanwhile, Dale and Meemaw go to the casino and Georgie makes a big mistake when Dale leaves him in charge of the store.

The ladies prepare for Joss' engagement party; Karen awaits her test results; April relies on Daniel to keep Lucy and her safe; Savi considers revealing her change of heart.

Residency Director Dr. Leanne Rorish (Marcia Gay Harden) puts her four new first year residents to work immediately on their first day at Angels Memorial Hospital, in the busiest, most notorious hospital ER in the nation.