In Victorian England, a coach meets with disaster, forever changing the fate of the accident's sole survivors.

Hostility grows between Jonathan and Dio, and forgotten secrets emerge; Jonathan is suspicious about the secret to the mask.

Jonathan and Speedwagon confront Dio, but they are caught off-guard when a reversal sends shockwaves through the family.

Jonathan meets Zeppeli, the master of Hamon energy, who trains him how to use Hamon to repair his body and deal devastating attacks; Dio recruits Jack the Ripper and sends him to kill Jonathan.

Jonathan and his fellow travelers face the devious Dio at Windknight's Lot, where an onslaught of evil ensues as Dio summons two undead warrior knights.

Jonathan's heroism transforms a foe's hatred into humanity; the crew is driven into a knightly training camp.