When publishing mogul Bradford Meade (Alan Dale) hands the reins of Mode, the bible of the fashion industry, over to this son, Daniel (Eric Mabius), he specifically hires Betty Suarez as his son’s new assistant - mostly because she's the only woman in NYC Daniel won't sleep with. Though this "player" is reluctant to accept her at first, Betty's indomitable spirit and bright ideas will eventually win him over. Neither of them really knows the ins and outs of the fashion world, but the two are a formidable team against those who will do anything to see them fall.
Betty unwittingly loses "the book," a proof of the entire magazine, complete with un-retouched photos of a popular actress (guest star Sarah Jones) who looks less than lovely without a little computer magic. If the photos get out, it could create huge embarrassment for Mode. Wilhelmina plots to use the snafu to bring Daniel down. Meanwhile, Bradford Meade secretively inquires about Fey Sommer's final effects.
It's Christmastime at Mode, as Betty questions her relationship with Walter and her feelings for Henry, the accountant. Daniel, on the other hand, is sure of his love for Sofia (Salma Hayek), despite her doubt, and Marc frantically tries to win over Wilhelmina in fear of losing his life. Meanwhile Amanda vies for Betty's position as Daniel's new assistant by planning and executing a perfect Mode holiday party that fills the office with Styrofoam snow and holiday wishes.
It's Thanksgiving Day, and everyone at Mode celebrates in their own special way. Betty is torn between work and home; Daniel attends his parents' brunch in hopes of seeing Sofia (guest star Salma Hayek); Marc and Amanda camp out at the Mode offices, and Wilhelmina attempts something she's never done before - cooking a turkey.
In an effort to boost their chances of getting pregnant, Molly encourages Mike to join her in giving up caffeine and eating healthier, causing them both to have major withdrawals.
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