Lola and Emily find themselves on opposing sides when the key witness in a murder trial is the victim’s 11-year-old daughter. Also, when Mark’s key witness in a case winds up in a Vegas jail, Luke accompanies Mark on a road trip to the desert to bring the witness back to testify.
Meredith takes a step forward with Will but quickly regrets it, while Maggie senses a disconnection with Andrew. Meanwhile, Amelia reevaluates her relationship with Owen, and Catherine comes to town and shakes things up with April and Jackson.
Meredith and Maggie support Amelia on an important day. Jo reveals she has been keeping secrets from Alex, while Arizona and Callie deal with the impact of their custody arrangement. Meanwhile, Ben leaps into action to help a friend.
Miranda continues to deal with the aftermath of Ben's reckless decisions and forms an advisory committee to recommend proper disciplinary action. Meanwhile, April and Jackson try to put their differences aside for the sake of their baby, and Arizona makes a decision that could affect her relationship with Callie forever.
James convinces a hockey player with a severe spinal injury to undergo risky surgery that could enable him to walk again. Also, James invites Walter's wife, FDA executive Julianna Wallace (Wendy Moniz), to Bunker Hill to persuade her to approve a new skin bio-printing trial.
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