When an art dealer is shot in his gallery, Castle and Beckett discover that a bullet is missing from the crime scene.
After notoriously murdering “The Gravedigger,” sniper-on-the-loose Jacob Broadsky (guest star Arnold Vosloo) strikes again. This time, his victim is a prior suspect who escaped the justice system. More determined than ever to catch Broadsky, Booth uses his own military training to beat the sniper at his own game. He vows to go to any lengths necessary to ensure that justice is served. In a heated and dramatic face-off, Booth has minutes to catch Broadsky before he hits an intended target, but Broadsky’s skill and speed may prove to be a worthy adversary. Meanwhile, Angela’s father (guest star Billy F. Gibbons) visits the Jeffersonian and insists on naming his unborn grandchild.
When the same tragedy occurs to people who don't appear connected, Danny must find what they have in common and who is responsible. Meanwhile, Frank helps an old friend come to terms with his alcoholism.
When an off-duty cop tries to stop a bank robbery, he is disarmed by the "Bonnie and Clyde" robbers who take his gun and wound an innocent bystander. Meanwhile, Frank considers getting romantically involved with a woman from the ACLU who happens to be Erin's acquaintance.
When a devoutly religious young woman is found murdered in a cemetery, Danny questions her family who is shocked to learn that she had a secret relationship.
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