Vice cop Casey Shraeger finds herself suddenly transferred into the NYPD Homicide Unit, where she begins to learn the secrets and eccentricities of her new co-workers.

One of New York's oldest family of criminals, the Boorlands, go on a crime spree that has the detectives running all over the city. When Walsh (Jeremy Renner) and Shraeger (Amber Tamblyn) learn the surprising reason behind the family's cluster of crimes, they use this information to negotiate its end. Meanwhile, Sergeant Brown (Terry Kinney) tells Shraeger that Walsh's former partner, Kowalski, was a dirty cop and asks her to dig into Walsh's past; and Cole's (Adam Goldberg) efforts to keep his criminal past a secret are threatened by the arrival of Frank Lutz (Ryan O'Nan), Cole's former partner in crime, on "The Unusuals."

Banks and Delahoy run into trouble on an undercover sting while Casey discovers that a personal favor throws a wrench into another case.

Beaumont tries to keep her financial woes a secret after she's caught in the middle of a pawn shop robbery.

Cole (Josh Close) receives a disturbing call from Lutz (Guest star Ryan O'Nan), who believes Walsh (Jeremy Renner) knows he killed Kowalski and is on his way to Walsh's diner. Lutz poses as a customer and strikes up a conversation with Walsh before pulling a gun and firing. Lutz escapes and Walsh discovers that one stray bullet went through the diner into his apartment, striking Beaumont. Cole fears for his future as Walsh and Casey begin to uncover pieces of his past relationship with Lutz. Meanwhile, Banks (Harold Perrineau) continues to be haunted by the number 42 while he and Delahoy (Adam Goldberg) investigate a string of bus robberies. One passenger, a clairvoyant, tries to help them catch the robber with her predictions, causing Banks to question whether fate can be stopped, on "The Unusuals."

Officer Lewis Powell fell off the wagon and woke up in an alley covered in blood with his off duty weapon missing. Lewis tells Walsh he may have killed someone the night before during an alcohol-induced blackout and asks for his help. While Walsh and Shraeger scramble to put together the pieces of Lewis' missing night, Banks and Delahoy learn they are both victims of identity theft. While investigating the murder of the man pretending to be Leo Banks, Delahoy and Banks peer into the lives of the men who have stolen their identities and consider changing the way they live their lives , on "The Unusuals."