When Buchio, a 20-year-old cat, is reborn as a nekomata, his family happily accepts him. As young Mutsumi worries about her missing father, Jiro the crow tengu comforts her. Tales of a mysterious yet warm day-to-day life shared by humans, gods, and yokai begin here.

As Buchio ponders what to do with his new life, Yuri the bakegitsune teaches him how to transform. Rain, a kappa in Ryo's class who has a crush on him, sees a certain someone for romantic advice...

Takumi and Buchio encounter a mythical beast said to bring misfortune to those who hear its voice. Mutsumi learns that Jiro once killed a giant serpent, but it turns out the serpent remained as a vengeful spirit...

Buchio starts attending driving school, where he meets a tsukumogami who's inhabited a car. Meanwhile, at the elementary school, a raiju falls from a rift in the sky...

Seeing Buchio enjoying a happy life with his family makes Yuri remember her tumultuous past with her own family, leaving her angry and irritable. Then she finds herself caught up in an unbelievable incident...

After returning from the parallel world, Yuri is finally ready to stop living in the past. To set herself free, she sets off to visit the family home she left behind many years ago under the crushing weight of pride and tradition.