Robin rediscovers a group of single girls who like to party; Ted pitches Barney a design for his company's headquarters.
Betty strives for a scoop with a new designer; Marc enlists clueless Amanda and Matt in his effort to sabotage Betty; Daniel attends bereavement sessions, where he meets Natalie (Jamie-Lynn Sigler), who takes him out for the night.
Wilhelmina makes Marc be Betty's temporary assistant; Betty decides to cover the worst jobs in New York; Hilda causes trouble for Archie's campaign; Daniel hires a beautiful but inept assistant.
Marc encourages Justin to befriend the cheerleaders at school; Bennett and Natalie lure Daniel deeper into the Community of the Phoenix; Wilhelmina tries to find money for Nico's blackmail payment.
Matt asks someone to be his "plus one" at Wilhelmina's fundraiser; Daniel hires Natalie to be his assistant, making Betty feel obsolete; Amanda keeps a secret from Betty; Daniel discovers that his mother has a history with Cal Hartley.