Vanessa Lachey stars as Special Agent in Charge of NCIS Pearl Harbor Jane Tennant, who, with her unwavering team of specialists, balances duty, family and country, while investigating high-stakes crimes involving military personnel, national security and the mysteries of the sun-drenched island paradise itself. In the premiere episode, An experimental Naval aircraft crashes on Oahu, and Tennant and her team must find who is behind it before classified state secrets are exposed.

Tennant and NCIS must take down a notorious team of international thieves using WWII explosives on armored trucks before they complete their final big job and disappear. Also, Kai searches for a new place to live; Lucy and Whistler have a standoff about their previous encounter; and Ernie's mission in the field is more alarming than he expected.

Kai goes undercover with one of Hawai'I's oldest surf gangs when a petty officer who's trying to help wayward kids find a new path in the Marines is murdered.

When a beloved Paniolo (Hawaiian cowboy) is shot while out riding his horse, Jane and her team must gain the trust of the Paniolo community to help find the culprits and protect the Paniolo's life. Also, Kai tries to convince his stubborn father to see a doctor.

When a Japanese sailor is killed on American soil and evidence links the case to the previous murder of the victim's girlfriend back in Japan, NCIS must find the murderer before the wrong person is accused and the case triggers a diplomatic crisis. Also, Captain Milius makes a personal request to Tennant.

When Tennant and NCIS are tasked with finding a kidnapped globe-trotting social media star, they find that she is not who her husband, or her followers, think she is. Also, the team is at odds with Whistler, who has an agenda of her own.