Taking place three months after the ending of Series Two, it’s now Spring 1939. Changes are happening at Skeldale, and everyone has to learn how to adjust. Tristan is now a qualified vet. James enters a new stage of his life with Helen and at the practice. After being made a joint business partner by Siegfried, James pushes to take on more responsibility via the Ministry of Agriculture’s new bovine TB testing scheme, but unexpected challenges risk pushing James to his breaking point. As a potential Second World War looms, the Skeldale family has to consider their purpose in Darrowby and beyond.