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When Apollo (Justin Long), a kind-hearted traveling performer and ingenious cricket, lands in the village of the Funny Little Bugs, the whole kingdom is about to be disrupted. The beloved reigning queen bee Marguerite (Kate Mara) is soon kidnapped on the orders of her jealous and treacherous cousin Wendy, who sets up a trap to frame Apollo for the crime, causing panic and despair in the hive.