In the season finale, while Mystic Falls presents a screening of "Gone with The Wind" in the town square, Damon's memories of Katherine in 1864 mix with the reality of Elena in the present day. Stefan pays a terrible personal price for his attempt to prevent a tragedy, and Sheriff Forbes (recurring guest star MARGUERITE MacINTYRE) makes a deadly mistake while trying to keep everyone safe. More than one life hangs in the balance as the consequences of the sacrifice ritual play out to a horrifying conclusion.
Damon is willing to go to extraordinary lengths to prevent Elena from being a victim of Klaus's (recurring guest star JOSEPH MORGAN) plan to break the curse. Damon's actions put him at terrible odds with Stefan and the tension that started brewing between the brothers over their differing opinions of how to help save Elena's life blows up into a full-scale conflict. Tyler returns to Mystic Falls after receiving a disturbing phone call. With the full moon marking the arrival of the sacrifice ritual, Stefan and Elena spend a romantic day together ... afraid it might be their last.
Stefan and Damon are furious when they discover that Elena has defied them both to make a new ally and take control of the plan to out-maneuver Klaus. Disagreement over how to protect Elena leads to growing tension between the Salvatore brothers. Meanwhile, Stefan has his hands full trying to keep a confused and frightened Jenna safe. Flashbacks to 1491 reveal Katherine’s introduction to Elijah and Klaus, along with the origin of the moonstone curse. Elena learns shocking new information about Klaus’ motives.
As the full moon arrives, Elena prepares for whatever Klaus (JOSEPH MORGAN) has planned and Tyler faces his second transformation. The terrifying events spin out of control.
In order to lure a vengeful vampire hunter named Rayna Cruz (recurring guest star LESLIE-ANNE HUFF) away from his friends in Mystic Falls, Stefan heads to New Orleans, where Valerie (recurring guest star ELIZABETH BLACKMORE) has learned of a safe house that can protect him. Once there, he comes face to face with Klaus Mikaelson (guest star/former series star JOSEPH MORGAN – The Originals), who soon becomes suspicious of his old friend's unexpected arrival in his city. Meanwhile, Enzo, who is working on behalf of a mysterious organization called The Armory, approaches Damon and Bonnie with a plan that could protect Stefan from Rayna. Finally, after learning that Alaric has decided to leave for Dallas with the twins immediately, Caroline offers to accompany them on the trip and finds herself contemplating her own future in Mystic Falls.
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