When Cybertron is threatened by a colossal black hole, Optimus Prime and the Autobots have no choice but to evacuate the entire planet. A mysterious visitor holds out one hope for salvation... but will a sneak attack by Megatron seal Cybertron's fate?

With the help of their human friends, the Autobots construct a new headquarters on Earth, but will Megatron discover their secret base before it's even finished?

Coby, Lori, and Bud try to help the Autobots maintain their disguises during their stay on Earth, but soon find that concealing their new friends is harder than they think. The real danger comes when an encounter with the Decepticons threatens to expose the Autobots!

Optimus sends Hot Shot and Red Alert to the Speed Planet to search for the first Cyber Planet Key, while Bud finds the Autobots' first clue to the location of the Omega Lock. In the battle for the ancient artifact, the veteran Landmine pushes himself to the limit and beyond. Will he be able to hold off the Decepticons long enough to recover the Omega Lock?

On the Speed Planet, Hot Shot has no choice but to disobey Optimus when Decepticon thugs attack the innocent. Meanwhile, back on Earth, Bud cajoles Scattorshot into taking him on a trip into space, but adventure turns to terror when a stray missile sends them hurtling to the Earth

As the Decepticons close in on the Autobots' Earth base, Hot Shot tries to learn more about the Cyber Planet Key from Clocker and Breakdown. They take him to one of Override's favorite racetracks, and Hot Shot finds himself in a high-speed contest with Megatron!