A civil war rages on the planet Cybertron between the heroic Autobots and the evil Decepticons. With the planet's resources depleted, the Autobots—led by their leader, Optimus Prime—set out on a mission to scout for new energy sources. But a surprise attack from Megatron and the Decepticons will leave both sides marooned and deactivated on the distant planet Earth for 4 million years! When they finally reawaken, the battle between good and evil continues on present-day Earth.

Using their robotic powers to transform, the Autobots and Decepticons disguise themselves as Earth vehicles, and a new chapter begins in their war between good and evil. Megatron and the Decepticons plan to capture Earth's energy by force to build a ship to take their newfound power back to Cybertron. It's up to the Autobots and their new human allies, Sparkplug and Spike, to defeat the Decepticons and save both worlds!

The battle for Earth's resources rages on! The Decepticons' ship is complete, and the odds are stacked against Optimus Prime, Bumblebee, Jazz and the others in their battle to stop them. When an Autobot trap is foiled by the cunning Megatron, all seems lost—but will the evil Starscream's hunger for power give the Autobots the edge they need to finally defeat the Decepticons? This is the thrilling conclusion to the original miniseries that started it all!

Presumed to be defeated, the Decepticons have returned! After receiving a message from Shockwave on Cybertron, Megatron comes out of hiding and orders the construction of a "Space Bridge" to transport Earth's energy to Cybertron. When Bumblebee and Spike uncover his plan they are captured, and Bumblebee is "reprogrammed" to lead the Autobots straight into a trap!

After his accidental transportation to Cybertron, Megatron returns—and he wants to get his hands on an antimatter formula created on Earth that could mean limitless power for the Decepticons! Bumblebee and Spike race to warn the Autobots, while their new human ally Chip attempts to hide the data. When Soundwave locates Chip and steals the formula, Megatron plots to use its power on the Autobots...

While protecting an Earth military plant from a Decepticon attack, Optimus Prime is severely wounded. His only hope for survival is a new Cosmotron unit. The good news is that Wheeljack's got one—the bad news is that it's in his lab back on Cybertron. Chip and an Autobot team must teleport across the Space Bridge and evade Megatron’s attacks in order to restore Optimus’s power before it's too late!